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Four taxa differing by size were described in these localities :
– H. concudense, MN 12,
– H truyolsi, MN 13 (the largest and the less represented),
– H. gromovae, MN 13,
– H. periafricanum, MN 13 (the smallest).
Their specific or subspecifc nature is not clear.
Dispersion diagrams of metapodials, first and second phalanges illustrate their size differences.