Concud, El Arquillo, Valdecebro II
Four taxa differing by size were described in these localities :
H. concudense, MN 12,
H truyolsi, MN 13 (the largest and the less represented),
H. gromovae, MN 13,
H. periafricanum, MN 13 (the smallest).
Their specific or subspecifc nature is not clear.
Dispersion diagrams of metapodials, first and second phalanges illustrate their size differences.